12042 Betta pugnax (Cantor, 1849) Osphronemidae Gouramies Macropodusinae Anabantiformes Teleostei Penang betta Asia: Malaysia, Singapore and Indonesia. ZMH 19602 Betta pugnax ZMH 967 Betta pugnax ZMH 18598 1894 Betta brederi ZMH 18599 1894 Betta brederi ZMH 19600 1894 Betta brederi ZMH 18597 1901 Betta brederi ZMH 19603 1901 Betta pugnax ZMH 19604 1901 Betta pugnax ZMH 18594 1902 Betta pugnax ZMH 4623 1959 Betta pugnax ZMH 4621 1971 Betta pugnax ZMH 4622 1971 Betta pugnax ZMH 4624 1971 Betta pugnax AMNH 57054 1985 Betta pugnax